David Watkins, who played rugby union and rugby league for Wales as well as for each sport's British and Irish Lions teams, has died at the age of 81.

Watkins starred at fly-half in union and stand-off in league.

The Rugby Football League said Watkins would "be remembered as one of the best cross-code converts from Wales".

The governing body added: "He became a key figure in a golden era for the Salford club, making more than 400 appearances for the Red Devils, and scoring almost 3,000 points as they were crowned champions twice in the space of three seasons in the mid-1970s."

Wales Rugby League said: "Dai, as he was affectionately known, is recognised as one of the greatest Welshmen to have ever pulled on a rugby shirt."

On social media, Newport RFC said: "We are extremely saddened to hear that David 'Dai' Watkins, one of the greatest players the club has ever produced, has passed away."

Watkins rose to prominence in Wales after joining Newport RFC in 1961, helping them famously beat New Zealand 3-0 in 1963, the year in which he made his Wales debut.

On the 1966 Lions tour to Australia and New Zealand, Watkins started all six Tests at fly-half and captained in two Tests.

In 1967, when moving to the professional ranks of rugby league at Salford for 16,000, Watkins was barred from playing union.

He went on to produce mesmerising performances for Salford, Wales and Great Britain before moving into coaching.

Watkins guided Great Britain to the 1977 World Cup final, in which they lost narrowly to Australia, and went on to coach Wales as well as help found Cardiff Blue Dragons league team.

He returned to union as Newport team manager in 1992 before becoming chairman, then president, a title he also held with regional team Dragons after they were formed in 2003.

Watkins' passing was announced shortly before Newport hosted Pontypool in the Welsh Premiership, with flags lowered to half mast in tribute.

Tributes also came from leading figures in both sports.

Ex-Wales and Great Britain forward Jim Mills posted on social media: "Heartbreaking news the legendary David (Dai) Watkins has passed away, a wonderful player in both codes of rugby, the only man to captain the Lions in RU & RL.

"Dai was such a lovely man and will be greatly missed. He will never be forgotten. My deepest sympathy to his wife Jane & family."

Regional team Dragons RFC, who are also based in Newport, wrote: "Our thoughts of us all @dragonsrugby are with everyone at @NewportRFC and the family of Dai Watkins."

The union Lions said Watkins was "a true legend of both codes".