Alex Dombrandt has been recalled to Steve Borthwick's initial 36-man England squad for Saturday's Calcutta Cup match against Scotland at Murrayfield.

The Harlequins number eight withdrew from the Six Nations opener with Ireland earlier this month through injury, but returned to club action against Saracens in the Premiership Cup on Saturday.

Alex Coles and Henry Pollock have also been added to the squad, with Curtis Langdon and Alex Lozowski dropping out.

Northampton lock Coles played for his club in the Premiership Cup on Friday night after recovering from a wrist injury he picked up in mid-January.

Tom Curry and George Martin are also back in training after picking up knocks during England's dramatic 26-25 win over France last weekend.

Borthwick's side lost their opening game in the competition 27-22 at reigning champions Ireland.